By 2012, Alex is a lieutenant and the Tactical Action Officer aboard the Arleigh Burke class flight I destroyer USS John Paul Jones, while Stone is the commanding officer of Arleigh Burke class flight IIA destroyer USS Sampson. Alex is also in a committed relationship with Samantha and wants to marry her, but is afraid to ask her father for permission. During the opening ceremony for the RIMPAC naval exercises, Alex brawls with Japanese officer Captain Nagata (Tadanobu Asano), the latest in a string of incidents that could result in his discharge at the end of RIMPAC. Meanwhile, Samantha, a physical therapist, accompanies retired Army veteran and amputee Mick Canales (Gregory D. Gadson) on a hike on Oahu to help him adapt to his prosthetic legs.
Five alien ships arrive in response to the NASA signal. One ship collides with an orbital satellite and crashes in Hong Kong, while four others land in the water near Hawaii. Arleigh Burkes Sampson, John Paul Jones, and the Japan Maritime Self-Defense Force Kongo class destroyer (which is a Japanese modification of the Arleigh Burke class) Myōkō investigate, but are trapped when one of the ships erects an impenetrable force field around the Hawaiian islands. The destroyers attempt to establish contact, but the aliens open fire: Sampson and Myōkō are destroyed, and John Paul Jones is damaged with the commanding and executive officers killed. Initially attached to Sampson, Alex, who was one of the few out to take a closer look at the alien vessels, now boards John Paul Jones and takes command as the most senior officer left on the ship. Although initially wanting to attack the alien ships in retaliation for his brother's death, Alex is convinced to break off and recover survivors from Myōkō, with Captain Nagata among them. This act prompts the aliens to call off another attack. Since the barrier prevents the use of radar and sonar, Nagata reveals that they can use tsunami warning buoys around Hawaii to track the alien vessels' movements.
During a night-time battle, the aliens and the humans exchange pot shots, with the Arleigh Burke class John Paul Jones sinking two alien ships but unable to lock onto the third. An alien soldier is recovered from the water; from the brief contact with the alien, the humans determine that the aliens are very sensitive to sunlight. Alex uses his ship to lure the alien vessel close to shore, where he and Nagata shoot out its bridge windows at sunrise, then proceed to fire at the alien vessel full force. Both ships are destroyed in the ensuing battle.
Because the ship that crashed in Hong Kong was their communications ship, the aliens take over the NASA array, using it to signal their home planet. Samantha and Mick avoid detection and run into a fleeing NASA scientist. Believing that a larger invasion might be imminent, the scientist acquires a radio which Samantha uses to warn Alex. He had planned to destroy the array with his ship, but since John Paul Jones has been destroyed, the survivors are forced to return to base and acquire the only available naval vessel, the Iowa class battleship USS Missouri. Although a museum ship, Alex and his crew are able to reactivate Missouri with the aid of the retired veterans preserving her. The battleship engages the alien ship and destroys its barrier but ends up with only one high-explosive shell remaining. Alex uses it to fire on the array, but before the defenseless Missouri can be attacked one more time, F/A-18 Hornet fighters from the RIMPAC fleet (which scrambled to attack as soon as the barrier fell) arrive and eliminate the alien ship. The NASA array explodes, destroying the aliens and their equipment.
Back on shore, the naval personnel, along with Mick, are honored for their actions in defeating the aliens. After the ceremony, Alex asks Admiral Shane for his daughter's hand in marriage; the admiral refuses, but invites him to talk about the matter over a chicken burrito.
In a post-credits scene set in rural Scotland, three schoolchildren and a handyman break open a car-sized object that has apparently fallen from the sky. They run off in terror after seeing an alien hand reach out and grab the opening.
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